Biological process: protein-membrane adaptor activity^{(2015, 2021, 2025)} identical protein binding^{(2005, 2019, 2025)} cytoskeleton-nuclear membrane anchor activity^{(2020, 2025)} lamin binding^{(2010, 2014, 2025)} nucleotide binding⁽²⁰²⁵⁾ chromatin binding^{(2019, 2025)} DNA binding^{(2019, 2025)} structural molecule activity⁽²⁰⁰⁵⁾ spindle pole body-nuclear membrane anchor activity⁽²⁰²⁰⁾ protein-transporting ATPase activity⁽²⁰²⁵⁾
± 4 protein-macromolecule adaptor activity²⁰²³ microtubule binding²⁰⁰² GTP binding²⁰²⁵ ATP binding²⁰²⁵

Molecular function: meiotic attachment of telomere to nuclear envelope^{2017, 2022, 2025} nuclear matrix anchoring at nuclear membrane^{2010, 2025} spermatogenesis^{2003, 2004, 2005, 2010, 2024, 2025} nucleus organization^{2012, 2015, 2019, 2025} protein localization to nuclear envelope^{2012, 2015, 2025} protein homooligomerization^{2015, 2025} protein heterooligomerization^{2015, 2025} centrosome localization^{2006, 2010, 2016, 2019, 2025} cell differentiation²⁰²⁵ nucleokinesis involved in cell motility in cerebral cortex radial glia guided migration^{2016, 2025}
± 40 positive regulation of cell migration^{2010, 2025} nuclear migration along microfilament^{2010, 2025} nuclear migration^{2002, 2005, 2007, 2025} meiotic cell cycle²⁰²⁵ homologous chromosome pairing at meiosis^{2013, 2015} spermatid development²⁰¹⁸ vulval development^{2006, 2025} response to mechanical stimulus^{2015, 2025} regulation of protein localization²⁰²⁴ protein-containing complex assembly²⁰²⁴ post-embryonic development^{2006, 2025} ossification^{2015, 2025} nuclear migration along microtubule^{2024, 2025} nuclear envelope organization^{2007, 2012} nervous system development^{2006, 2025} mitotic sister chromatid segregation^{2019, 2025} meiotic telomere clustering²⁰¹³ locomotion^{2006, 2025} egg-laying behavior^{2006, 2025} centrosome cycle^{2019, 2025} centromere clustering at the mitotic interphase nuclear envelope^{2012, 2020} cell division²⁰²⁵ telomere tethering at nuclear periphery²⁰¹³ subtelomeric heterochromatin formation²⁰²¹ spindle pole body duplication²⁰¹³ regulation of establishment or maintenance of cell polarity regulating cell shape²⁰¹⁷ regulation of centrosome duplication²⁰⁰⁹ regulation of cell migration²⁰⁰⁵ protein transmembrane transport²⁰²⁵ nucleus localization²⁰¹⁹ nuclear migration involved in conjugation with cellular fusion²⁰¹³ mitotic spindle pole body insertion into the nuclear envelope²⁰¹⁸ mitotic spindle organization²⁰⁰² mitotic sister chromatid cohesion²⁰¹³ meiotic attachment of telomeric heterochromatin to spindle pole body²⁰¹⁹ karyogamy²⁰¹³ intracellular protein transport²⁰²⁵ intracellular protein localization²⁰⁰⁶ establishment of mitotic sister chromatid cohesion²⁰¹³ embryo development ending in birth or egg hatching²⁰⁰⁶

Cellular component: nuclear envelope^{2002, 2006, 2007, 2009, 2010, 2012, 2013, 2017, 2019, 2021, 2022, 2024, 2025} meiotic nuclear membrane microtubule tethering complex^{2006, 2010, 2024, 2025} nuclear inner membrane^{2004, 2007, 2010, 2017, 2019, 2025} membrane^{2007, 2012, 2022, 2025} nuclear membrane^{2024, 2025} phragmoplast^{2015, 2021, 2025} endoplasmic reticulum membrane^{2015, 2025} chromosome, telomeric region^{2007, 2020, 2022, 2025} condensed nuclear chromosome^{2007, 2025} sperm head-tail coupling apparatus^{2018, 2024, 2025}
± 30 nucleus^{2002, 2018, 2025} cytoplasm^{2002, 2025} cytoskeleton²⁰²⁵ spindle pole body^{2013, 2025} nuclear periphery^{2012, 2013, 2018} microtubule^{2005, 2025} endoplasmic reticulum^{2014, 2021} perinuclear region of cytoplasm^{2007, 2025} nucleoplasm^{2019, 2025} nuclear outer membrane^{2019, 2025} mitotic spindle pole body^{2009, 2016} endosome membrane²⁰²⁵ centrosome^{2010, 2025} acrosomal membrane²⁰¹⁰ Golgi apparatus²⁰²⁵ spindle microtubule²⁰¹⁰ spindle²⁰²¹ site of double-strand break²⁰¹⁴ outer acrosomal membrane²⁰²⁵ old mitotic spindle pole body²⁰¹⁸ new mitotic spindle pole body²⁰¹⁸ microtubule organizing center²⁰²⁵ meiotic spindle pole body²⁰²⁰ meiotic spindle pole²⁰²¹ interphase microtubule organizing center²⁰¹⁴ inner plaque of mitotic spindle pole body²⁰¹⁸ half bridge of spindle pole body²⁰¹³ cytoplasmic vesicle²⁰²⁵ chloroplast outer membrane²⁰²⁵ Sad1-Kms1 LINC complex²⁰¹⁹